Hi,

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tilesize=4000

this means the max. size of the tiles is 4000x4000,
together with the setting - maxcubesize=2200 - only 1 tile per cube face will be generated,

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levels=1

only 1 resolution level will be generated (no multi-resolution)

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maxsize=8000

the maximum size of a spherical/cylindrical image,
a larger file will be downscaled to that size,

when a cubical pano (or a converted cubical pano) was used this setting will be ignored at the maxcubesize setting will be used,

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maxcubesize=2200

the maximum size for a cubeface side (2200x2200)
larger images will be downscaled,

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jpegsubsamp=422

the used jpeg color sub-sampling setting,
see here for details - http://en.wikipedia.org/wiki/Chroma_subsampling

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jpegoptimize=true

the jpeg compression uses two kinds of compression - first a lossy quantizing, and than a lossless compression of the quantized data, and second part, the lossless compression was done by a huffman compression,
and this huffman compression uses internally a table of codes for compression,

by default this codes are predefined which results in a good but not optimal compression,
to get the best possible compression this codes can be optimized by analyzing the complete quantized data of the image,
but to be able to analyze the complete image the whole image must be stored in memory,
without optimizing only memory for a few lines of the images is needed and the coding is faster,

normally optimizing isn't a problem, but for very large (e.g. gigapixel panos) and few system memory, the storing of the images for code analyzing can be a memory problem,

btw - have also a look into the "kmakemultires.config",
there the settings are also explained,

best regards,
Klaus